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Notes 8.1
Species Interactions
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Directions
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Introduction
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Species
Interactions
- Populations
- Population
Dynamics
- Habitats
and Niches
- Population
Distribution
- Competition
- Symbiosis
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Populations
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Population
- Definition:
A population is a group of individuals of a particular species among
which breeding is possible
- All the members of
a species in a locale. Ex.
All the blue mussels Mytilus
edulis on the rocky shores of Palos Verdes California or all
the horseshoe crabs Limulus polyphemus of Delaware Bay
Individual
- Definition:
An individual is a single organism
- A single member of
a species. Ex.
A blue mussel Mytilus edulis or a horseshoe
crab Limulus polyphemus
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Population
Dynamics
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Population Dynamics
- Definition:
Population dynamics are the changes in populations over time.
- Dynamic changes include
population growth and decline.
- Populations can also
be stable over time.
- Definition:
Stable is constant or unchanging.
Population Growth
- Definition:
Population growth is an increase in the numbers of individuals.
- Growth will occur
if the numbers of births and immigrants exceeds the numbers of deaths
and emigrants.
- Definition:
Immigrants are
individuals entering a population by moving in from elsewhere
- Definition:
Emmigrants are individuals leaving a population by moving eleswhere
Population Decline
- Definition:
Population growth
is a decrease in the numbers of individuals.
- Decline will occur
if the numbers of deaths and emigrants exceeds the numbers of births
and immigrants.
Population Models
- Definition:
Population models
are mathematical formulae that mimic changes in natural populations.
- Exponential growth
is a model that mimics growth without limits which occurs in nature
when populations are young and have colonized environments that are
ideal for their growth or when some limiting factor in the environment
has been eliminated.
Population Models
- Definition:
Population models
are mathematical formulae that mimic changes in natural populations.
Exponential Growth
Model
- Exponential growth
is a model that mimics growth without limits
- The increase in
the number of individuals over a given period of time is equal to
the birth rate minus the death rate multiplied by the current number
of individuals
- The number of individuals
graphed against time yeilds a J-shaped curve
- Growth without
limits occurs in nature when populations are young and are located
in environments that are ideal for their growth or when some factor
limiting their growth has been eliminated
Logistic Growth Model
- Logistic growth is
a model of self-limiting growth
- The increase in the
number of individuals over a given period of time is equal to the
birth rate minus the death rate multiplied by the carrying capacity
minus the carrying capacity divided by the current number of individuals
- Definition:
Carrying capacity is the maximum number of individuals that the environment
can support.
- The number of individuals
graphed against time yeilds a S-shaped curve
- Self-limiting growth
occurs in nature when populations are older and are located in environments
that include factors that resist growth
Population Regulation
- Definition:
Population regulation
is the control of the number of individuals in a population.
- Regulation is brought
about by environmental factors that determine the carrying capacity.
It occurs when the severity of environmental resistance is directly
related to the number of individuals in the population. Such
factors are density dependent.
- Competition for a
limited resource, degradation of the environment by metabolic wastes,
attacks by predators, and diseases are examples of density dependent
environmental factors that determine the carrying capacity.
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Habitats
and Niches
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Environmental Challenges
- Various environmental
factors require special efforts by organisms to survive and prosper
- Environmental factors
are either abiotic or biotic
- Definition:
Abiotic is nonliving (ex. temperature, salinity, pH, light, pressure,
dissolved oxygen, currents, tides, waves, exposure to air)
- Definition:
Biotic is living (ex. competition, predation, symbiosis)
Habitat
- Definition:
The kind of site with respect to biotic and abiotic factors where
a species is typically found.
- For example, a coastal
ecosystem is divided into rocky shore, mud flat, and oyster reef habitats
- The habitat can be
envisioned as the home address of a species and refers to the environment
Microhabitats
- Definition:
A microhabitat is a subdivision of a habitat with a unique set of
abiotic and biotic factors.
- Describes small scale
variations in habitats
Microhabitats vs.
Habitats
- Habitats contain many
microhabitats occupied by different species.
- Microhabitats contain
fewer species. For example, the rocky shore habitat is subdivided
into upper surfaces of rocks, under rocks, and tidal pools microhabitats
Ecological Niche
- Definition:
An ecological niche is the position of a species in nature defined
by all its requirements for survival including food, shelter, competition,
predation etc.
- The niche can be envisioned
as the ecological definition of a species and refers to the living
organisms
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Population
Distribution
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Range of Habitats
- The range of habitats
and microhabitats occupied by a species is determined by its ability
to deal with biotic pressures and to tolerate abiotic extremes
- Each species is able
ot tolerate certain environmental conditions and thus will be found
where those conditions prevail
Abiotic Gradients
- Definition: An
abiotic gradient is a gradual change in an abiotic factor over
a distance
- Occurs when one environment
blends into another
- Examples: temperature
with latitude, salinity with distance from river mouth, pressure with
depth

Range of Toleration
- Definition:
The range of toleration is the range of values of a particular factor
within which members of a population can survive.

Optimum Condition
- Definition:
The optimum condition is a value of an Abiotic Factor at which the
individuals of a population operate best.
- An ideal measure
- If values vary from
optimum the health of an organism declines
- Populations have their
areas of greatest abundance at positions along a gradient that are
close to the optimum condition
Stressful Conditions
- Values of an Abiotic
Factor sufficiently distant from the optimum condition to cause a
strain on the organism
Limits of Tolerance
- Definition:
Limits of tolerance are values of
an Abiotic Factor at which the individuals of a population are dying
- Cannot be exceeded
without the complete loss of the population
- Populations have their
areas of scarcity at positions along a gradient that are close to
their limits of tolerance
Effects of
Gradients on Species Distributions
- Species have different
ranges of toleration with different optimum conditions and limits
of tolerance
- Species ranges of
toleration are found in different positions along an environmental
gradient
- Each species has its
own area of greatest abundance where it experiences its optimum conditions
- Each species has its
own areas of scarcity where it experiences environmental stress
Ranges of Toleration
Vary Among Species
- Some species have
narrow ranges of tolerance while others have wide ones
- Steno- = narrow
- Eury- = wide
- Thermal (heat),
haline (salt), baric (pressure), bathic (depth), topic (place)
- Stenothermal = tolerates
only a narrow range of temperature
- Euryhaline = tolerates
a wide range of salinity
Geographical Range
- Definition:
A geographical range is the overall geographical area occupied by
a population or a species.
- Spatial distribution
of species on a regional or global scale
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Competition
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Definition of Competition
- Definition:
Competition is a struggle for resources that are renewable but limited.
- Definition:
Renewable is continuously supplied.
- Definition:
Limited is not
enough to go around.
- Examples: food, space,
light (plants), etc.
Types of Competition
- Definition:
Intraspecific competition is between members of same species
- Definition: Interspecific
competition is between
members of different species
Competitive Exclusion
- Definition:
Competitive exclusion the rejection of a species from an area due
to its inability to compete with others
- According to Gause's
Principle no two species that occupy the same ecological niche can
coexist in the same location for long, one will competitively exclude
the other.
- Competitive exclusion
will occur if one of the competing species produces enough individuals
to prevent the population of the other species from expanding into
its habitat.
- Competitive exclusion
will also occur if one competing species invades the habitat of the
other species and successfully eliminates it by monopolizing the limited
resources
Competitive Coexistance
- Two species compete
but both continue to exist
- The niches of the
competing species overlap but do not totally coincide
Resource Partioning
- Definition: Resource partitioning is an adaptive process
by which species divide up resources in order to avoid competition.
- Acheived when species specialize and use different portions of
a habitat.
- Allows for competitive coexistance
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Symbiosis
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Definition of Symbiosis
- Definition:
Symbiosis is two species living together in a close relationship that
has evolved over a long period of time.
- Definition:
Symbiont is the partner that always benefits from the relationship
and is usually the smaller of the two.
- Definition:
Host is the partner that may or may not benefit from the relationship
and is usually the larger of the two.
Types of Symbiosis
- Mutualism
- Commensalism
- Parasitism
- Endosymbiosis
Mutualism
- Definition:
Both species (symbiont and host) benefit from the partnership

Examples of Mutualism
- Cleaner Fish (symbionts)
pick parasites off Larger Reef Fish (hosts) that come to cleaners
to be groomed
- Anemones (symbionts)
live on and are carried about on the shells of Hermit Crabs (hosts)
which are protected from predators by the anemones stingers
- Clown Fish (symbionts)
live with and are protected by Anemones (hosts) which they in turn
feed
- Zooxanthellae (symbionts)
live inside and nourishes Coral Polyps (hosts) which provide them
with nutrients
- Algae (symbionts)
live inside and nourishes Fungi (hosts) which provide them with nutrients
(Lichens)
Commensalism
- Definition:
One species benefits (symbiont) and the other (host) is neither benefited
nor harmed by the partnership

Examples of Commensalism
- Wedding Shrimp (symbionts)
live protected inside the cavity of Glass Sponges (hosts)
- Gobies and Pea Crabs
(symbionts) live inside the protection of the burrows of Inkeeper
Worms (hosts)
Parasitism
- Definition:
One species is benefited (parasite symbiont) and the other is harmed
(host)
Examples of Parasitism
- Fish Lice (symbionts)
suck the blood of Fish (hosts)
- Liver Flukes (symbionts)
live inside and consumes the livers of Seals (hosts)
- Tapeworms (symbionts)
live in the gut and saps nourishment from Fish (hosts)
Endosymbiosis
- Definition:
One species (endosymbiont) lives inside another (host)
Examples of Endosymbiosis
- Zooxanthellae (endosymbionts)
live inside and nourishes Coral Polyps (hosts) which provide them
with nutrients
- Algae (endosymbionts)
live inside and nourishes Fungi (hosts) which provide them with nutrients
(Lichens)
- Liver Flukes (endosymbionts)
live inside and consumes the livers of Seals (hosts)
- Tapeworms (endosymbionts)
live in the gut and saps nourishment from Fish (hosts)
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