Holoplankton
are species of invertebrates that spend their entire lives from hatching
until death as plankton.
Some species
of invertebrates have their embryonic development in the plankton and
become planktonic adults. These species spend their entire lives in the
plankton. They are the most abundant type of zooplankton.
Holoplanktonic
Crustacea
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Copepods are tiny
holoplanktonic crustaceans. Copepods make up over 90% of the zooplankton
by number and are the most abundant marine animals.
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Observe the photograph
of a copepod.
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Observations:
- Describe the anatomy
of the copepod.
- Describe the features
that help it stay afloat and swim.
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Holoplanktonic
Crustacean Larvae
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Nauplii are the first
planktonic, larval stages in the development of many crustacea,
including copepods. They hatch from eggs and begin to feed and grow
immediately. They grow and develop into copepodites which ultimately
develop into adult copepods.
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Experimental
Set Up:
- We photographed
copepod nauplii, copepodites, and adult copepods from a zooplankton
sample taken off the coast of Southern California.
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Observe
the photographs of nauplii, copepodites and adult copepods.
Nauplii |
Copepodites |
Adult
Copepods |
 |
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|
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 |
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Observations:
- How do the nauplii
differ from the copepodites and adult copepods you just observed?
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Holoplanktonic
Copepod Diversity
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Copepods
are diverse as well as abundant. A typical zooplankton sample will
contain several different species.
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Experimental
Set Up:
- We photographed
several species of adult copepods from a zooplankton sample taken
off the coast of Southern California.
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Observe
the photographs of adult copepod species.
Cyclops |
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Calanus |
 |
Eurytemora |
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Observations:
- How do the adult
copepods you just observed differ? How are they similar?
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Holoplanktonic
Predator
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Arrow worms are holoplanktonic
invertebrates which feed on copepods. They are fast swimmers and
can easily capture their prey.
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Experimental
Set Up:
- We obtained a
preserved specimen of an arrow worm, placed it in a watchglass
filled with water, and photographed it using a dissecting microscope.
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Observe
the photographs of the arrow worm.
Whole
Worm |
Anterior
End |
Head |
 |
 |
 |
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Observations:
- Describe the special
structures near its mouth that are used to impale copepods.
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Gelatinous Holoplankton
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There are several
types of holoplanktonic species whose bodies are jelly-like. They
are called gelatinous zooplankton. These forms include medusae,
siphonophores, salps and larvaceans.
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Experimental
Set Up:
- We photographed
several species of gelatinous zooplankton from a zooplankton sample
taken off the coast of Southern California.
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Observe
the photographs of gelatinous zooplankton species.
Medusae |
 |
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Siphonophores |
 |
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Salps |
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Larvaceans |
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Observations:
- Describe the differences
between medusae, siphonophores, salps, and larvaceans.
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Additional Gelatinous
Plankton
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Gelatinous zooplankton
include medusae, ctenophores and siphonophores.
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Experimental
Set Up:
- We
photographed several additional species of gelatinous zooplankton
obtained from various locations.
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Observe
the photographs of gelatinous zooplankton.
Medusae |
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Ctenophores |
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Siphonophores |
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Observations:
- Describe the features
that gelatinous zooplankton all share in common.
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Meroplankton
are the planktonic larvae of nektonic or benthic species.
Some species
of invertebrates spend their adult lives on or in the benthos but their
embryos develop in the plankton. Some species of fishes spend their adult
lives as nekton but their eggs and embryos develop in the plankton. These
planktonic larvae act as a dispersal stage being distributed by currents
to new sites where they settle to the bottom and grow into adults.
Meroplanktonic
Crustacean Nauplii
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Barnacles live cemented
to hard substrates as adults but their larval stages are meroplanktonic.
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Experimental
Set Up:
- We obtained a
prepared slide of a barnacle nauplius and photographed it at high
power under a compound microscope.
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Observe
the photographs of the nauplii. |
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Observations:
- How does a nauplius
larvae differ from the adult?
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Meroplanktonic
Echinoderm Larvae
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Starfish live on
the benthos as adults but their embryos develop in the plankton.
The embryo passes through two distinct stages before its final metamorphosis.
The bipinnaria larva grows into a brachiolaria larva which is ideally
suited to life in the plankton. These meroplanktonic larvae undergo
dramatic changes as they metamorphose.
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Experimental
Set Up:
- We obtained prepared
slides of bipinnaria and brachiolaria larvae. We photographed
both larval forms at high power under a compound microscope.
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Observe
the photograph of the bipinnaria larva. |
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Observe
the photograph of the brachiolaria larva. |
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Observations:
- What does the
bipinnaria have that the brachiolaria does not have?
- How does this
feature benefit the bipinnaria for its temporary life in the plankton?
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Meroplanktonic
Larvae
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Many meroplanktonic
larvae look very different from the adult forms they develop into.
Many were misidentified by marine biologists who described them
as different species. These mistakes were corrected once developmental
sequences were clarified.
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Experimental
Set Up:
- We photographed
several different meroplanktonic larvae from a zooplankton sample
taken off the coast of Southern California.
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Observe
the photographs of the meroplanktonic larvae.
Type
of Larva Form |
Photograph |
Adult
Form |
Veliger |
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1. |
Trochophore |
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2.
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Zoea |
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3.
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Bipinnaria |
 |
4.
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Brachiolaria |
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5. |
Nauplius |
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6.
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Fry |
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7.
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Observations:
- Use your books
to help you identify the adult forms that produce the meroplanktonic
larvae in the photographs above.
- You can print
out this chart and add the names of the adults or make a similar
chart in your notebook.
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Demersal
plankton are species that live in the sea bottom during the day but rise
up into the plankton at night.
Demersal species are temporary
members of the plankton and are taken by plankton nets in shallow waters
throughout the night.
Demersal Crustacea
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Amphipods
are crustaceans which may be planktonic or benthic. Some species are
demersal plankton. |

Observe the photograph
of an amphipod.
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Observations:
- Describe the features
of the amphipod that make it capable of living temporarily in
the plankton.
- Describe the features
of the amphipod that make it capable of living in the benthos.
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